Development is remarkably reproducible, producing organs with the same size, shape, and function repeatedly from individual to individual. Yet, these reproducible organs are composed of highly variable cells. My laboratory focuses on the mechanisms that produce diversity in cell size in the Arabidopsis leaf and sepal (outermost green leaf-like floral organ) and the mechanisms that ensure the same size and shape leaves and sepals are produced from these variably sized cells. Today I will tell you three short stories about the variable organ size2 (vos2) mutant and how it disrupts the reproducibility of sepal sizes by disrupting translation, about the jaw-d mutant and how it disrupts leaf flatness, and about the asymmetric division of the moss apical cell and how it generates the spiral pattern of leaves (phyllids).