This series of lectures seeks to inject fresh energy into debates about England’s plural and protracted Reformations by adopting the concept of generation as its analytical framework. Its aim is to investigate how the tumultuous religious developments of the sixteenth and seventeenth centuries not merely transformed the generations that experienced it, but also reconfigured the nexus between memory, history, and time. The lectures examine how age and ancestry were implicated in the theological and cultural upheavals of the era and explore how the Reformation shaped the horizontal relationships that early modern people formed with their siblings, kin, and peers, as well as the vertical ones that tied them to their dead ancestors and their future heirs. They highlight the important part that the family has played in shaping our knowledge of the Reformation past and in the making of its archive. They contend that religious revolution had both biological and social dimensions.